Human cDC1 enhance cytotoxic function of CD226+ terminally exhausted tumor-infiltrating lymphocytes.

Publication date: Dec 01, 2025

Prevention or reversal of T cell exhaustion is a major objective of cancer immunotherapy. However, few models exist to generate, characterize and modulate exhausted human T cells, particularly within solid tumors in vivo, which likely hampers the discovery and translation of novel therapeutics. In this study we describe a humanized mouse model where functional human CD8+ T cells specific for the tumor antigen NY-ESO-1 develop in vivo from human CD34+ hematopoietic stem cells genetically modified to express a HLA-A *0201-restricted NY-ESO-1 specific T cell receptor (TCR). HLA-A *0201+ NY-ESO-1+ expressing A375 melanoma tumors engrafted in these mice and were refractory to treatment with anti-PD-1 despite being infiltrated with NY-ESO-1 specific T cells. Tumor-Infiltrating Lymphocytes (TIL) upregulated tissue resident memory (TRM) markers CD103 and CD69 along with exhaustion markers PD-1, TIGIT, and CD39 relative to T cells from other organs. Further, TILs failed to secrete cytokines TNF and IFNγ following in vitro stimulation with conventional Type I Dendritic Cells (cDC1), indicative of terminal exhaustion. However, cDC1 stimulation of the terminally exhausted NY-ESO-1 specific TILs led to enhanced tumor killing that was associated with increased CD107a and Granzyme B expression that was restricted to a subset of CD226+ NY-ESO-1 specific TILs. These findings establish a novel platform to investigate T cell exhaustion in human tumors and suggest a role for cDC1 in enhancing terminally exhausted TIL cytotoxic function.

Concepts Keywords
Cd103 Animals
Hampers Antigens, Differentiation, T-Lymphocyte
Therapeutics Antigens, Differentiation, T-Lymphocyte
Tumor Antigens, Neoplasm
Vivo Antigens, Neoplasm
CD226
CD8-Positive T-Lymphocytes
cDC1 dendritic cells
Cell Line, Tumor
Cytotoxicity, Immunologic
Dendritic Cells
Humanized mice
Humans
Lymphocytes, Tumor-Infiltrating
Melanoma
melanoma
Mice
T cell exhaustion
THBD protein, human
Thrombomodulin
Thrombomodulin

Semantics

Type Source Name
disease MESH tumor
disease MESH T cell exhaustion
pathway REACTOME Translation
drug DRUGBANK Esomeprazole
disease MESH melanoma
pathway KEGG Melanoma

Original Article

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